1. Key Laboratory of the Zoological Systematics and Evolution, Institute of Zoology Chinese Academy of Sciences Beijing China
2. Department of Biology, Faculty of Natural and Computational Sciences Assosa University Assosa Ethiopia
3. University of Chinese Academy of Sciences Beijing China
4. College of Sciences Bahir Dar University Ethiopia
5. Center for Macroecology, Evolution and Climate, Natural History Museum of Denmark University of Copenhagen Copenhagen Denmark
6. Center for Excellence in Animal Evolution and Genetics Chinese Academy of Sciences Kunming China