1. Centre for Bone and Arthritis Research, Institute of Medicine, Sahlgrenska Academy, University of Gothenburg, Gothenburg, Sweden
2. Department of Anatomy and Cell Biology, University of Oulu, Oulu, Finland
3. Department of Functional Genomics, Institut de Génétique et de Biologie Moléculaire et Cellulaire (IGBMC), Université de Strasbourg (UdS), Collège de France, Illkirch, France