1. Natural History Museum University of Oslo PO Box 1172 Blindern 0318 Oslo Norway
2. EA7316 Université de Rennes I 263 Av. du Général Leclerc 35042 Rennes Cedex France
3. Department of Biosciences Center for Ecological and Evolutionary Synthesis (CEES) University of Oslo PO Box 1066 Blindern 0316 Oslo Norway