1. Centre for Image AnalysisUppsala University Uppsala 75124 Sweden
2. Department of Pharmaceutical BiosciencesUppsala University Uppsala 75124 Sweden
3. Faculty of Medicine and Life SciencesUniversity of Tampere Tampere 33014 Finland
4. Faculty of Biomedical Sciences and EngineeringTampere University of Technology Tampere 33720 Finland
5. BioImage Informatics Facility of SciLifeLab Uppsala 75124 Sweden