1. Vaccine and Infectious Disease Organization (VIDO), Home of the International Vaccine Centre (InterVac), University of Saskatchewan; Saskatoon Saskatchewan S7N 5E3 Canada
2. Nantes-Atlantic College of Veterinary Medicine and Food Sciences and Engineering, UMR BioEpAR, LUNAM Université, Oniris; Nantes F-44307 France
3. Epidemiology and Risk Analysis in Animal Health, CS 40706, INRA, UMR1300 Biology; Nantes F-44307 France