1. Montana Cooperative Wildlife Research Unit Wildlife Biology Program W.A. Franke College of Forestry and Conservation University of Montana Natural Sciences Room 205 Missoula Montana 59812 USA
2. Wildlife Biology Program, W.A. Franke College of Forestry and Conservation University of Montana 32 Campus Drive Missoula Montana 59812 USA
3. Idaho Department of Fish and Game 2885 West Kathleen Avenue Coeur d’Alene Idaho 83815 USA
4. U.S. Geological Survey Montana Cooperative Wildlife Research Unit Natural Sciences Room 205 University of Montana Missoula Montana 59812 USA
5. Panthera and Wildlife Biology Program, W.A. Franke College of Forestry and Conservation University of Montana Natural Sciences Room 205 Missoula Montana 59812 USA