1. Department of Ecology, College of Urban and Environmental Sciences, and Key Laboratory for Earth Surface Processes of the Ministry of EducationPeking University Beijing China
2. Key Laboratory of Adaptation and Evolution of Plateau Biota, Northwest Institute of Plateau BiologyChinese Academy of Sciences Xining China
3. Department of Earth, Atmospheric, and Planetary SciencesPurdue University West Lafayette Indianapolis USA
4. Department of AgronomyPurdue University West Lafayette Indianapolis USA