1. a Division of Genetics and Development Krembil Research Institute, University Health Network, Toronto, Ontario, Canada
2. b Department of Orthopaedic Surgery Keio University School of Medicine, Tokyo, Japan
3. c New World Laboratories, Laval, Quebec, Canada
4. d Institute of Medical Sciences, University of Toronto, Toronto, Ontario, Canada
5. e Department of Surgery and Spine Program University of Toronto, Toronto, Ontario, Canada
6. f Electron Microscope Laboratory Keio University School of Medicine, Tokyo, Japan