1. a Department of Molecular Biology and Genetics, Johns Hopkins University School of Medicine, Baltimore, Maryland, USA
2. b Department of Ophthalmology, Wilmer Eye Institute, Johns Hopkins University School of Medicine, Baltimore, Maryland, USA
3. c Shiley Eye Institute, University of California, San Diego, La Jolla, California, USA
4. d The Solomon H. Snyder Department of Neuroscience, Johns Hopkins University School of Medicine, Baltimore, Maryland, USA
5. e Institute of Genetic Medicine, Johns Hopkins University School of Medicine, Baltimore, Maryland, USA