1. MRC Human Immunology Unit Weatherall Institute of Molecular Medicine and University of Oxford, NIHR Biomedical Research Centre Oxford England UK
2. Section of Musculoskeletal Disease University of Leeds Leeds UK
3. Department of Clinical Immunology Oxford University Hospitals NHS Trust Oxford UK
4. Division of Rheumatology Immunology and Allergy Department of Medicine Brigham and Women's Hospital Harvard Medical School Boston MA USA