1. Department of Applied Ecology North Carolina State University Raleigh North Carolina 27607 USA
2. Washington Department of Fish and Wildlife Olympia Washington 98504 USA
3. National Park Service Olympic National Park Port Angeles Washington 98362 USA
4. U.S. Geological Survey Forest and Rangeland Ecosystem Science Center Port Angeles Washington 98362 USA
5. California Department of Fish and Wildlife Redding California 96001 USA