1. Research Unit of Plant Breeding and Plant Genetic Resources Conservation, National Institute of Agricultural ResearchRegional Agricultural Research Center of Meknes Meknes 50100 Morocco
2. Laboratory of Plant Biotechnology and Molecular Biology, Department of Biology, Faculty of ScienceUniversity Moulay Ismail Meknes 50100 Morocco
3. Instituto de Agricultura SostenibleConsejo Superior de Investigaciones Científicas (CSIC) Córdoba E‐14080 Spain