1. Lancaster Environment Centre; Lancaster University; Lancaster LA1 4YQ UK
2. School of Natural Sciences and Psychology; Liverpool John Moores University; Liverpool L3 3AF UK
3. School of Physical and Geographical Sciences; Keele University; Keele ST5 5BG UK
4. School of Geography; Queen Mary University of London; London E1 4NS UK
5. School of Biological and Chemical Sciences; Queen Mary University of London; London E1 4NS UK