1. Department of Biology, Faculty of Sciences Ferdowsi University of Mashhad Mashhad Iran
2. Department of Invasion Ecology, Institute of Botany The Czech Academy of Sciences Průhonice Czech Republic
3. Department of Range and Watershed Management, Faculty of Natural Resources and Environment Ferdowsi University of Mashhad Mashhad Iran
4. Department of Botany, Research Center for Plant Sciences Ferdowsi University of Mashhad Mashhad Iran