1. Research Center for Bioscience and Technology; Tottori University; Yonago Japan
2. Japan Society for the Promotion of Science; Tokyo Japan
3. Chromosome Engineering Research Center; Tottori University; Yonago Japan
4. Division of Human Genome Science; Department of Molecular and Cellular Biology; School of Life Sciences; Faculty of Medicine; Tottori University; Yonago Japan
5. Department of Biomedical Science; Institute of Regenerative Medicine and Biofunction; Graduate School of Medical Sciences; Tottori University; Yonago Japan
6. Central Institute for Experimental Animals; Kawasaki Japan
7. Cancer Research UK Cambridge Institute; Li Ka Shing Centre; University of Cambridge; UK