1. Department of Agricultural, Food and Nutritional Science; University of Alberta; 116 Street and 85 Avenue, Edmonton Alberta T6G 2P5 Canada
2. Department of Biochemistry; University of Alberta; 116 Street and 85 Avenue, Edmonton Alberta T6G 2H7 Canada
3. French Associates Institute for Agriculture and Biotechnology of Drylands, The Jacob Blaustein Institutes for Desert Research; Ben-Gurion University of the Negev, Sede Boqer Campus, Midreshet Ben-Gurion; 8499000 Israel