1. Resource Ecology Group Wageningen University Droevendaalsesteeg 3a Wageningen 6708 PB The Netherlands
2. School of Life Sciences University of KwaZulu‐Natal Westville Campus Durban 4000 South Africa
3. Department of Terrestrial Ecology Netherlands Institute of Ecology (NIOO‐KNAW) Droevendaalsesteeg 10 Wageningen 6708 PB The Netherlands