1. U.S. Department of Agriculture, Agriculture Research Service, Warmwater Aquaculture Research Unit Stoneville Mississippi USA
2. Thad Cochran National Warmwater Aquaculture Center, Mississippi Agriculture and Forestry Experiment Station, Department of Wildlife, Fisheries, and Aquaculture, College of Forest Resources Mississippi State University Stoneville Mississippi USA
3. Thad Cochran National Warmwater Aquaculture Center, Aquatic Research and Diagnostic Laboratory, Department of Pathobiology and Population Medicine, College of Veterinary Medicine Mississippi State University Stoneville Mississippi USA
4. Department of Medicine and Epidemiology, School of Veterinary Medicine University of California–Davis Davis California USA