1. Department of Ecology and Evolutionary Biology University of Toronto Toronto ON M5S 3B2 Canada
2. Carl Woese Institute for Genomic Biology University of Illinois at Champaign‐Urbana Urbana IL 61801 USA
3. Research School of Biology Australian National University Canberra ACT 2601 Australia
4. Koffler Scientific Reserve University of Toronto King ON L7B 1K5 Canada