1. School of Biological Sciences; Nanyang Technological University, 60 Nanyang Drive; Singapore 637551 Singapore
2. Department of Biological Sciences; National University of Singapore, 14 Science Drive 4; Singapore 117543 Singapore
3. Department of Mathematics; National University of Singapore, 10 Lower Kent Ridge Road; Singapore 119076 Singapore
4. Yau Mathematical Sciences Center, Tsinghua University, Haidian District; Beijing 100084 China