1. Nashville VA Medical Center; Nashville Tennessee USA
2. Departments of Surgery and Cell and Developmental Biology and the Epithelial Biology Center; Vanderbilt University School of Medicine; Nashville Tennessee USA
3. Division of Gastroenterology, Department of Medicine; Vanderbilt University School of Medicine; Nashville Tennessee USA
4. Department of Pathology; Vanderbilt University School of Medicine; Nashville Tennessee USA
5. Department of Cancer Biology; Vanderbilt University School of Medicine; Nashville Tennessee USA