1. Department of Biological Sciences Columbia University New York NY 10027 USA
2. Department of Biochemistry and Molecular Biophysics Columbia University New York NY 10032 USA
3. Department of Pathology, Microbiology and Immunology Vanderbilt University School of Medicine Nashville TN 37235 USA
4. Department of Biochemistry Vanderbilt University School of Medicine Nashville TN 37235 USA
5. Department of Chemistry Vanderbilt University Nashville TN 37235 USA