1. IRD, UMR 234, GET; Toulouse France
2. CNRS, GET; Toulouse France
3. UPS (OMP), GET, Université de Toulouse; Toulouse France
4. Now at St Anthony Falls Laboratory; University of Minnesota; Minneapolis Minnesota USA
5. Aix Marseille Université, IRD, CNRS, CEREGE UM34, Aix-en-Provence, France