1. Department of Developmental Biology and Neurosciences, Graduate School of Life Sciences; Tohoku University; Aobayama Aoba-ku Sendai 980-8578 Japan
2. Frontier Research Institute for Interdisciplinary Sciences; Tohoku University; Aobayama Aoba-ku Sendai 980-8578 Japan
3. Department of Biochemistry and Molecular Biology, Faculty of Agriculture and Life Science; Hirosaki University; Hirosaki 036-8561 Japan
4. Division of Biological Science, Graduate School of Science; Nagoya University; Furo-Cho, Chikusa-ku Nagoya 464-8602 Japan