1. Jodrell Laboratory; Royal Botanic Gardens; Kew; Richmond Surrey TW9 3DS U.K.
2. Unit of Ecology & Evolution; Department of Biology; University of Fribourg; 1700 Switzerland
3. Institute of Forest Genetics; Federal Research and Training Centre for Forests; Natural Hazards, and Landscape; Hauptstrasse 7 Vienna 1140 Austria
4. Identification Systems and Morphometrics Section; Herbarium; Royal Botanic Gardens; Kew; Richmond Surrey TW9 3AB U.K.