1. Department of Zoology University of Otago Dunedin New Zealand
2. School of Biological, Earth and Environmental Sciences (BEES) University of New South Wales Sydney Australia
3. Institute for Biodiversity and Environmental Research Universiti Brunei Darussalam Gadong Brunei Darussalam
4. Faculty of Science Universiti Brunei Darussalam Gadong Brunei Darussalam