1. INRA; UMR 1331 Toxalim INRA-INP; 180 chemin de Tournefeuille, BP 93173; F31027; Toulouse cedex 3; France
2. INRA; UMR 1331 Toxalim INRA-INP; 400 route des Chappes, BP 167; F06903; Sophia Antipolis cedex; France
3. LUNAM, Oniris; USC 1329 INRA, LABERCA; Atlanpôle-La Chantrerie, BP 50707; F44307; Nantes; France