1. Department of Biology and Chemistry Changwon National University Changwon Republic of Korea
2. Department of Biology, School of Arts and Sciences National University of Mongolia Ulaanbaatar Mongolia
3. Laboratory of Vegetation Ecology and Economy, Botanic Garden and Research Institute Mongolian Academy of Sciences Ulaanbaatar Mongolia
4. Key Laboratory of Plant Resources Conservation and Sustainable Utilization, South China Botanical Garden Chinese Academy of Sciences Guangzhou China