1. Department of Plant Sciences Quaid‐ i- Azam University Islamabad 45320 Pakistan
2. State Key Laboratory of Systematic and Evolutionary Botany Institute of Botany Chinese Academy of Sciences Beijing 100093 China
3. University of Chinese Academy of Sciences Beijing 100049 China
4. CAS Key Laboratory of Mountain Ecological Restoration and Bioresource Utilization Chengdu Institute of Biology Chinese Academy of Sciences Chengdu Sichuan 610041 China