1. Center of Plant Ecology, Core Botanical Gardens Chinese Academy of Sciences Guangzhou China
2. Guangdong Provincial Key Laboratory of Applied Botany, South China Botanical Garden Chinese Academy of Sciences Guangzhou China
3. Environmental Horticulture Research Institute Guangdong Academy of Agricultural Sciences Guangzhou China
4. Key Lab of Ornamental Plant Germplasm Innovation and Utilization Guangzhou China
5. University of Chinese Academy of Sciences Beijing China
6. Guangzhou Institute of Forestry and Landscape Architecture Guangzhou China