1. Department of Life Science and Biotechnology, Faculty of Life and Environmental Science; Shimane University; Matsue Shimane Japan
2. Present address: Department of Biotechnology, Faculty of Engineering; Toyama Prefectural University; Imizu Toyama Japan
3. Present address: School of Chemical Engineering and Pharmacy; Wuhan Institute of Technology; Wuhan China
4. Graduate School of Science and Technology; Kumamoto University; Kumamoto Japan