1. Centre for African Conservation Ecology, Department of Zoology Nelson Mandela University Port Elizabeth South Africa
2. African Primate Initiative for Ecology and Speciation (APIES), Earth Stewardship Science Research Institute Nelson Mandela University Port Elizabeth South Africa
3. APIES, Department of Zoology and Entomology University of Fort Hare Alice South Africa
4. Department of Botany and Zoology Stellenbosch University Matieland South Africa
5. EPHE PSL Paris Université Paris France
6. UMR CNRS 6282 Biogéosciences Université de Bourgogne Dijon France