1. Sydney Brenner Institute for Molecular Bioscience, Faculty of Health Sciences University of the Witwatersrand Johannesburg South Africa
2. Division of Human Genetics, National Health Laboratory Service and School of Pathology, Faculty of Health Sciences University of the Witwatersrand Johannesburg South Africa
3. Childhood Complex Disease Genomics Section Center for Precision Health Research, National Human Genome Research Institute, National Institutes of Health Bethesda Maryland USA