1. Graduate School of Life SciencesHokkaido UniversityKita‐ku Sapporo060‐0810 Japan
2. Department of Marine Biodiversity ResearchJapanese Agency for Marine‐Earth Science and TechnologyYokosuka237‐0061 Japan
3. Creative Research InstitutionHokkaido UniversityKita‐ku Sapporo001‐0021 Japan
4. Precursory Research for Embryonic Science and TechnologyJapan Science and Technology AgencySaitama332‐0012 Japan