1. Biogéosciences, UMR CNRS 6282; Université de Bourgogne; 6 Boulevard Gabriel 21000 Dijon France
2. Laboratoire LBBE, UMR CNRS 5558; Université Claude Bernard Lyon 1; bâtiment Mendel, 43 boulevard du 11 novembre 1918 69622 Villeurbanne Cedex France
3. Department of Zoology; University of Venda; Private Bag X5050 Thohoyandou 0950 South Africa
4. Department of Integrative Biology and Evolution; Konrad-Lorenz-Institute of Ethology; University of Veterinarian Medicine Vienna; Savoyenstr. 1a A-1160 Vienna Austria