1. Division of Biological Sciences and Wildlife Biology Program University of Montana 32 Campus Drive Missoula Montana 59812 USA
2. Denali National Park and Preserve P.O. Box 9 Denali Park Alaska 99755 USA
3. Central Alaska Network Inventory and Monitoring Program 4175 Geist Road Fairbanks Alaska 99709 USA
4. Department of Mathematical Sciences University of Montana 32 Campus Drive Missoula Montana 59812 USA