1. Institut Pasteur, Unit of Molecular Prevention and Therapy of Human Diseases, Department of Infection and Epidemiology; rue du Dr. Roux F-75015 Paris France
2. CNRS, URA3012; rue du Dr. Roux F-75015 Paris France
3. Present address: CNRS, UMR7138; Quai Saint-Bernard F-75005 Paris France