1. Laboratoire d'Ecologie Microbienne, UMR1418 INRA, UMR5557 CNRS, UCBL, VetAgroSupUniversité de Lyon Villeurbanne 691002 France
2. Institute of Environmental Technology, Vietnam Academy of Science and Technology Hanoi Vietnam
3. iEES‐Paris (Sorbonne Université, UPEC, IRD, CNRS, INRA) Case 237, 4 Place Jussieu Paris Cedex 05 75005 France
4. Géosciences Environnement Toulouse (GET)Université de Toulouse, IRD, CNRS, UPS Toulouse 314005 France
5. IRD, UMR 242Institute of Ecology and Environmental Sciences – Paris (iEES‐Paris), c/o Soils and Fertilizers Research Institute (SFRI), Dong Ngac, Tu Liem Hanoi Vietnam
6. IRD, UMR 242Institute of Ecology and Environmental Sciences – Paris (iEES‐Paris) 32 Henri Varagnat Bondy 75005 France
7. IRD, Ban Sisangvone Vientiane BP 5992 Laos