1. Key Laboratory of Tibetan Medicine Research; Northwest Institute of Plateau Biology; Chinese Academy of Sciences; 23 Xin'ning Road Xining 810008 P. R. China
2. University of Chinese Academy of Sciences; Beijing 100049 P. R. China
3. Center of Mitochondria and Healthy Aging; College of Life Sciences; Yantai University; 30 Qingquan Road Yantai 264005 P. R. China
4. State Key Laboratory of Plateau Ecology and Agriculture; Qinghai University; Xining 810008 P. R. China