1. Graduate School of Biosphere Science; Hiroshima University; 1-7-1 Kagamiyama Higashi-Hiroshima 739-8521 Japan
2. Graduate School of Bioscience; Nagahama Institute for Bioscience and Technology; 1266 Tamura-cho Nagahama-shi Shiga 526-0829 Japan
3. Graduate School of Integrated Arts and Sciences; Hiroshima University; 1-7-1 Kagamiyama Higashi-Hiroshima 739-8521 Japan