1. School of Biological Sciences Washington State University Pullman, Washington 99164‐4236 USA
2. Royal Botanic Gardens, Kew Richmond, Surrey TW9 3AE UK
3. National Herbarium of New South Wales, Botanic Gardens of Sydney Australian Botanic Garden, Locked Bag 6002 Mount Annan 2567 New South Wales Australia
4. Evolution and Ecology Research Centre, School of Biological, Earth, and Environmental Sciences University of New South Wales Sydney Kensington NSW 2052 Australia
5. Centre for Ecology, Evolution and Behaviour, Department of Biological Sciences, School of Life Sciences and the Environment Royal Holloway University of London Egham TW20 0EX UK