1. Hubei Key Laboratory of Edible Wild Plants Conservation and Utilization Hubei Normal University Huangshi Hubei province 435002 China
2. Hubei Engineering Research Center of Special Wild Vegetables Breeding and Comprehensive Utilization Technology Hubei Normal University Huangshi, Hubei province 435002 China
3. National Experimental Teaching Demonstration Center for Biology Hubei Normal University Huangshi Hubei province 435002 China