1. School of Chemical Engineering & Technology China University of Mining and Technology Xuzhou Jiangsu 221116 P. R. China
2. Key Laboratory of Systems Microbial Biotechnology Tianjin Institute of Industrial Biotechnology Chinese Academy of Sciences Tianjin 300308 P. R. China
3. CAS Key Laboratory of Microbial Physiological and Metabolic Engineering State Key Laboratory of Microbial Resources Institute of Microbiology Chinese Academy of Sciences Beijing 100101 P. R. China
4. College of Life Sciences University of Chinese Academy of Sciences Beijing 100149 P. R. China