1. Boyce Thompson Institute; Ithaca NY USA
2. Plant Biology Section; Cornell University; Ithaca NY USA
3. Ministry of Education Key Laboratory of Biodiversity; Sciences and Ecological Engineering and Collaborative Innovation Center for Genetics and Development; Institute of Plant Biology; School of Life Sciences; Fudan University; Shanghai China
4. Department of Biology and the Huck Institutes of the Life Sciences; the Pennsylvania State University; University Park PA USA