1. Department of Fisheries and Aquatic Sciences, School of Fisheries and Wildlife, College of Agriculture and Environmental Sciences Bahir Dar University, Bahir, Dar Ethiopia
2. Department of Zoological Sciences Addis Ababa University Addis Ababa Ethiopia
3. Department of Biology College of Science Bahir Dar University, Bahir, Dar Ethiopia
4. Bahir Dar Fisheries and Other Aquatic Life Research Centre, Bahir, Dar Ethiopia