1. Department of NeurosciencePerelman School of Medicine at the University of PennsylvaniaPhiladelphia PA
2. Department of Clinical Studies–New Bolton CenterUniversity of Pennsylvania School of Veterinary MedicineKennett Square PA
3. Department of NeurologyPerelman School of Medicine at the University of PennsylvaniaPhiladelphia PA
4. Institut d'Investigacions Biomèdiques August Pi i Sunyer (IDIBAPS) and Institució Catalana de Recerca i Estuis Avançats (ICREA), Hospital Clínic, University of BarcelonaBarcelona Spain