1. Hebei Key Laboratory of Soil Ecology, Centre for Agricultural Resources Research, Institute of Genetics and Developmental Biology Chinese Academy of Sciences Shijiazhuang China
2. Laboratory of Molecular Iron Metabolism, The Key Laboratory of Animal Physiology, Biochemistry and Molecular Biology of Hebei Province, College of Life Science Hebei Normal University Shijiazhuang China
3. Institute of Genetics and Developmental Biology, State Key Laboratory of Plant Genomics Chinese Academy of Sciences Beijing China
4. Department of Biology Sonoma State University Rohnert Park California USA
5. University of Chinese Academy of Sciences Beijing China