1. Department of Gastroenterology and Metabolism, Graduate School of Biomedical and Health ScienceHiroshima University Hiroshima Japan
2. Research Center for Hepatology and GastroenterologyHiroshima University Hiroshima Japan
3. Natural Science Center for Basic Research and DevelopmentHiroshima University Hiroshima Japan
4. Center for Medical Specialist Graduate Education and Research, Institute of Biomedical and Health SciencesHiroshima University Hiroshima Japan