1. Department of Natural Resources and Environmental Science, Program in Ecology, Evolution, and Conservation Biology; University of Nevada Reno; Mail Stop 186 Reno NV 89557 USA
2. Department of Wildlife, Fisheries, and Conservation Biology; University of Maine; 5755 Nutting Hall Room 210 Orono ME 04469 USA
3. Washington Department of Fish and Wildlife; 2315 North Discovery Place Spokane Valley WA 99216 USA
4. Nevada Department of Wildlife; 6980 Sierra Center Parkway, Suite 120 Reno NV 89511 USA