1. Department of Natural History Sciences Graduate School of Science Hokkaido University Sapporo Japan
2. Frontiers in Environmental Sciences Graduate School of Agriculture Hokkaido University Sapporo Japan
3. Laboratory of Evolutionary Zoology and Genetics Federal Scientific Center of the East Asia Terrestrial Biodiversity Far Eastern Branch of the Russian Academy of Sciences Vladivostok Russia
4. Department of Natural History Sciences Faculty of Science Hokkaido University Sapporo Japan